Date of Acceptance: 27/12/2015 Acknowledgements We would like to thank all scientists and crew from the Autonomous Ecological Surveys of the Abyss (AESA) 2012 cruise, Alice Horton for assistance with the experiment at sea, Ben Boorman for design and building of apparatus and Maureen Procée and Gillian Martin for technical assistance and tuition in the laboratory. This work was carried out as part of the SERPENT project www.serpentproject.com. DJM was part-funded by NERC, UK (NE/G014744/1). BT received funding from the Rural and Environmental Science and Analytical Services Division, UK (RESAS).
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